An al-Qaeda leader in Yemen wanted in connection with the 2000 bombing of the American warship USS Cole has been killed in an air strike, officials say.
Witnesses say he was hit by a missile as he stepped out of a vehicle, along with another al-Qaeda militant, in a remote mountain valley in the southern province of Shabwa.
The strike, which was reportedly carried out by the CIA with the authorisation of the Yemeni government, came after an extended surveillance operation.
The US had offered a $5m (£3.1m) reward for information leading to Quso's capture or death.
A US official welcomed the death of the "senior terrorist operative". He told AFP that Quso had been planning further attacks against the US and Yemen.
The USS Cole was attacked in October 2000 while it was
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